THIS WEEK at HILTON POND
15-21 October 2007

Installment #376---
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BYE-BYE, RUBY-THROATS.
2007 WAS A VERY GOOD YEAR!

In spring, folks who enjoy hummingbirds get very enthusiastic and hang their feeders. When only a few hummers come, some of these same folks begin to wonder what's wrong. Usually, nothing is, it's just that people remember swarms of hummers they had in late summer and forget in spring there are NEVER many birds. After all, in very early April in the Carolina Piedmont there are only adult males, followed shortly thereafter in migration by adult females, and it takes a while before they all establish territories, mate, nest, and fledge enough young to drive numbers up a little. This whole process may take six weeks or so, meaning that even if your hummers arrive in mid-April it will be June before the earliest young start to fledge--and that's in the SOUTHERN states. So our usual response to all those frantic phone calls and E-mails is for folks to be patient, to wait until August for the usual hummingbird frenzy, and then to contact us in October AFTER the hummers are gone to ask whether this year's population was smaller than usual. Only then can we look at our data and tell you what our banding results say: The ruby-throats may have gone bye-bye, but 2007 was a very good year.

All text & photos © Hilton Pond Center

We started banding Ruby-throated Hummingbirds, Archilochus colubris, at Hilton Pond Center in 1984, mostly because we wanted to know how many we had at our feeders. Now it's 24 years later and all we can say is we learn many new things about ruby-throats every year; it follows that the more we learn about hummingbirds the more we realize there's still a lot we DON'T know. However, through banding--both at Hilton Pond and in Costa Rica on the ruby-throats' wintering grounds--we at least are discovering a few things about longevity, migration, and site fidelity. This week we'd like to share some of this info by summarizing our hummingbird banding results for 2007.

Chart 1: Click on chart above to open a larger image in a new browser window

The season started on 16 April, more than two weeks AFTER our early banding record of 27 March 1991; most years we band our first Ruby-throated Hummingbird (RTHU) at Hilton Pond Center on or before 10 April. Progress was excruciatingly slow with only five total bandings through 23 May, but things took off and we banded 23 birds by that month's end. For the rest of the spring and summer banding proceeded at a near-record pace (see Chart 1 above), finally tapering off in mid-September. The final capture occurred on 4 October, roughly our average late date for the last RTHU of the season.

Chart 2: Click on chart above to open a larger image in a new browser window

Although Chart 1 with all its squiggly lines shows nicely how banding progressed through the 2007 season, it's a bit easier to see how successful we were in by looking at the bar graph in Chart 2 just above. As shown, our best year ever was two years ago (2005) when we banded 226 RTHU. For some reason there was a big drop last year with only 164 hummers captured, but the Hilton Pond population--which includes both breeding birds and spring and fall migrants--rebounded nicely in 2007 to our third-best total of 204 birds. That was also well above our 18-year average of 168, a figure that does not include the six partial field seasons for which we were away from Hilton Pond for significant portions of the summer. If we can trust the red trend line on the chart above, Ruby-throated Hummingbirds do indeed appear to be on the increase at the Center; we doubt the upward curve is a result of our becoming more experienced at catching hummers, nor does it reflect more hours of trapping and mist netting.

Table 1: RUBY-THROATED HUMMINGBIRDS
BANDED AT
HILTON POND IN 2007, COMPARED TO ALL-TIME HIGHS AND
18-YEAR AVERAGES
Adult
Males
Adult
Females
Hatch- Year
Males
Hatch- Year
Females
TOTAL
2007
Totals
21
47*
71
65
204
Record
High
34
47*
108
68
226
18-year
Average
21
30
71
48
168
* Previous record for adult females was 45, set in 1998.

Note: We have been banding RTHU at Hilton Pond Center since 1984--a 24-year span--but for six of those years we were away for substantial periods during the summer; we do not include those years when calculating averages.

At the end of each hummingbird banding season, it's always interesting to look at the breakdown among age/sex classes as illustrated in Table 1 (above). Most years adult females outnumber adult males by about a 3:2 margin, while the ratio flips among young birds so that fledgling males outnumber young females. The adult ratios were a little skewed in 2007, primarily because we banded a record-high number of adult females (47) while capturing an exactly average number of adult males (21). Comparatively, juvenile RTHU were right on with almost exactly 60% of banded hatch-year birds being males. We caught an average number of young males (71), but the 65 young females were well above the 18-year average and almost set a new record. Had young males been banded at the same rate as young females, we might have surpassed our record of 226 total RTHU banded in a season at Hilton Pond Center.

It's hard to know whether the relatively poor showing by young males is simply a one-year blip or if there may have been something in the environment that cut their numbers. We doubt it was the Easter Freeze--which occurred before the majority of RTHU were even nesting this year--but it's worth noting male Ruby-throated Hummingbirds ARE smaller than females and might be more susceptible to inclement weather. The male also tends to get into more trouble as he goes about his business of establishing a territory and defending it against other males.

Even though 204 RTHU banded at Hilton Pond Center in 2007 didn't reach our all-time high of 226, we still encountered more hummers than usual--mostly because of an amazing influx of "old" birds from previous seasons. When we first reported on our 2007 hummingbird banding progress back on 31 May, we mentioned we already had recaptured 33 "old" hummers banded prior to this year. From 1984 through 2006, we had 23 returns on average per year, so we were elated in 2007 to end up recapturing 49 RTHU we banded in 2006 or before--an incredible 36% increase over the previous record of 36 set in 2000! Those 49 individuals are listed in Table 2 below, showing the banding date and when it was recaptured in 2007, as well as its age at banding and recapture and any previous returns. (We won't apologize that the list is long; in fact, we're excited it is!)

Table 2: 49 BANDED RUBY-THROATED HUMMINGBIRDS
RETURNING TO
HILTON POND IN 2007
(through 21 October)
(HY = hatch year bird, which becomes SY or 2nd year on 1 Jan. An adult bird whose hatching year is not known is called AHY, or "after hatch year"; on 1 Jan an AHY bird becomes ASY or "after 2nd year," and so on.)
Band
#
Recapture
Date
(2007)
Banding
Date
Age/Sex
at
Banding
Age
at
Recapture
Years of
Previous
Returns
Y15914
04/16
07/27/06
HY-M
2nd year
-
Y15920
04/23
08/07/06
HY-M
2nd year
-
Y15713
04/29
08/15/05
HY-F
3rd year
06
Y15998
04/29
09/13/06
HY-M
2nd year
-
Y15780
04/29
09/13/05
AHY-F
after 3rd year
06
Y15877
05/21
06/06/06
AHY-F
after 2nd year
-
Y15882
05/21
06/30/06
AHY-F
after 2nd year
-
Y15929
05/21
08/11/06
HY-M
2nd year
-
Y15712
05/22
08/11/05
AHY-F
after 3rd year
06
Y15770
05/22
09/01/05
HY-F
3rd year
06
Y15903
05/22
07/23/06
AHY-F
after 2nd year
-
Y16001
05/22
09/15/06
HY-M
2nd year
-
Y15875
05/22
06/05/06
AHY-F
after 2nd year
-
Y15695
05/24
08/01/05
HY-F
3rd year
06
Y15902
05/24
07/23/06
HY-M
2nd year
-
Y15131
05/24
07/11/02
AHY-M
after 6th year
03,04,05,06
Y15925
05/24
08/08/06
AHY-M
after 2nd year
-
Y15868
05/25
05/31/06
AHY-F
after 2nd year
-
Y15468
05/25
07/04/04
AHY-F
after 4th year
05,06
Y15981
05/25
09/06/06
HY-M
2nd year
-
Y15669
05/25
07/16/05
AHY-F
after 3rd year
none
Y15190
05/26
08/30/02
HY-M
6th year
03,04,05,06
Y15935
05/26
08/16/06
HY-F
2nd year
-
Y15939
05/26
08/17/06
AHY-F
after 2nd year
-
Y15869
05/28
05/31/06
AHY-F
after 2nd year
-
Y15917
05/28
07/30/06
HY-F
2nd year
-
Y15725
05/30
08/18/05
HY-F
3rd year
06
Y15332
05/30
07/23/03
AHY-F
after 5th year
05,06
Y15972
05/30
08/31/06
HY-F
2nd year
-
Y15311
05/30
06/08/03
AHY-F
after 5th year
04,06
Y15893
05/30
07/17/06
HY-F
2nd year
-
Y15682
05/30
07/26/05
HY-F
3rd year
06
Y15459
05/30
06/02/04
AHY-M
after 4th year
05
Y15948
06/04
08/23/06
HY-M
2nd year
-
Y15885
06/04
07/07/06
HY-F
2nd year
-
Y15894
06/05
07/17/06
HY-F
2nd year
-
Y15960
06/07
08/27/06
AHY-F
after 2nd year
-
Y15898
06/07
07/22/06
AHY-F
after 2nd year
-
Y16003
06/08
09/15/06
HY-M
2nd year
-
Y15895
06/09
07/17/06
AHY-F
after 2nd year
-
Y15685
06/10
07/27/05
AHY-F
after 3rd year
06
Y15582
06/14
08/30/04
HY-M
4th year
none
Y15689
06/17
07/28/05
AHY-F
after 3rd year
06
Y15969
07/16
08/30/06
HY-M
2nd year
-
Y15673
07/18
08/31/05
AHY-F
after 2nd year
none
Y15257
07/23
09/17/02
HY-F
6th year
NONE
Y15358
07/23
08/14/03
HY-F
5th year
04,05
Y15904
07/30
07/23/06
AHY-F
after 2nd year
-
Y15983
08/28
09/06/06
HY-M
2nd year
-

Banding hummingbirds provides interesting data, but RECAPTURING them tells us much more--even if the birds aren't encountered long distances away from the banding site. Thus, we'd like to point out some things in Table 2 (above) that should be interesting to the hummingbird enthusiast, especially when we toss in the additional results in Table 3 (below).

Table 3: RETURNS OF RUBY-THROATED HUMMINGBIRDS BANDED at HILTON POND,
1985-2007
(Does not include birds banded in 2007
that won't have their first return until next year)
AGE/SEX
Total
Returns
Total
Banded
Percent
Returns
Adult Males
53
425
12.5%
Adult Females
117
593
19.7%
Hatch-year Males
104
1,438
7.2%
Hatch-year Females
136
954
14.3%
All Age/Sex Classes
410
3,410
12.0%

The initial items of interest pertain to RTHU we first encountered at Hilton Pond Center last year, in 2006:

  • 29 of our 2007 returns were from 2006 (when we banded a mere 164 birds) for a return rate of 17.7%. This is an all-time high rate of return for hummers banded the preceding year. The old record was 17.0% set in 1992 from 130 RTHU banded in 1991. The average next-year returns since 1985 has been 10.5%.
  • Making up this year's 29 first-year returns from 2006 were 12 males and 17 females. With one exception, ALL these males were young birds hatched in 2006; this means 14.2% of the 77 young males banded last year returned in 2007--nearly TWICE our 24-year average of 7.2% for this age/sex class.
  • Of the 17 females banded last year that returned in 2007, only six were "young" hummers fledged in 2006; this gives a return rate of 13.0% on a total of 46 young females banded last year--only a only a little below their 24-year average of 14.3%. However, a whopping 35.5% of our 31 "old" females banded last year showed up again in 2007--nearly DOUBLING their 19.7% average.

Other gleanings from Table 2 above:

  • In 2007 we had 11 returns from 2005, and three each from 2002, 2003, and 2004.
  • 2007 is the fourth year we've had returning RTHU banded five years previously; the birds from 2002 make this the first time we've had THREE from that long ago.
  • Of this year's 49 returns, only two were originally captured in May, with five in June, 17 in July and again in August, and eight in September. These numbers make sense when we report the vast majority of our RTHU at Hilton Pond Center are banded from mid-July through early September. (We've never had a bird return after being banded in October--or even after 17 September for that matter.)

Perhaps the most interesting return for 2007 at Hilton Pond Center was Y15257, re-trapped on 23 July. Banded as a hatch-year bird in 2002, this sixth-year female was not seen again until this year--the first time in 24 years we've had a hummer go that long before recapture. Where she's been in the interim we'll never know. (Curiously, two hours after trapping the ancient and venerable Y15257 we pulled the string on Y15358, a fifth-year female from 2003 that returned in 2004 and 2005 but was not seen last year.)

On the other end of the scale for site fidelity are two males--Y15131 and Y15190; they were first caught in 2002, the former as an adult male that year and the latter as a fledgling. Both these birds have been back EVERY year since. As an after-sixth-year male, Y15131 ties with a female as the OLDEST Ruby-throated Hummingbirds known for Hilton Pond Center; that female was banded in 1997 and not seen after 2002, so Y15131 still has a chance to catch the record-holding nine-year-old RTHU documented by the federal Bird Banding Laboratory.

What does it mean when record numbers of "old" birds returned to Hilton Pond Center in 2007? We're not sure, exactly, but perhaps a mild hurricane season in the Gulf of Mexico last fall allowed for greater survival during southward migration. In any case, it's nice to see these familiar band numbers--especially when we consider that old Y15131 and Y15190 have been back and forth across the Gulf more times than we have for our Costa Rica banding expeditions!

In conclusion, we've shown you a bunch of numbers this week about new hummingbirds and those from years past, but that's about the only way we know to present information summarizing our annual hummer banding results. With our third-highest total of new birds (204) PLUS a record-setting 49 returns, we handled 253 Ruby-throated Hummingbirds at Hilton Pond Center in 2007, making it without doubt a very good year!

All text & photos © Hilton Pond Center

Gorget photos above represent the four different age/sex classes of
Ruby-throated Hummingbirds. (Can you figure them out?)
Below is a photo of the underside of a ruby-throat's tail. (Do you know its age and sex?)


Comments or questions about this week's installment?
Please send an E-mail message to INFO.

Be sure to scroll down for an account of all
birds banded or recaptured during the period,
plus other nature notes of interest.


Thanks to the following fine folks for recent gifts in support of Hilton Pond Center for Piedmont Natural History and/or Operation RubyThroat: The Hummingbird Project. Your tax-deductible contributions allow us to continue writing, photographing, and sharing "This Week at Hilton Pond." (Please see Support if you'd like to make a gift of your own.)

  • Elizabeth Boyd (on-line donation via Network for Good)
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  • South Carolina Wildlife Federation (Guided Field Trip fee)

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"This Week at Hilton Pond" is written & photographed
by Bill Hilton Jr., executive director of
Hilton Pond Center for Piedmont Natural History.

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BIRDS BANDED THIS WEEK at
HILTON POND CENTER

15-21 October 2007

SPECIES BANDED THIS WEEK:
Yellow-rumped Warbler--50

American Goldfinch--5
Carolina Chickadee--2
Palm Warbler--1
*
Pine Warbler--1
Chipping Sparrow--3
Northern Cardinal--7
Gray Catbird--1
Scarlet Tanager--3
Downy Woodpecker--2
White-throated Sparrow--7
Swainson's Thrush--1
Eastern Bluebird--1
House Finch--7
Brown-headed Cowbird--1
Gray-cheeked Thrush--1 *
Eastern Towhee--3
Brown Thrasher--1

American Robin--5
Common Grackle--1
*

* = New species for 2007


WEEKLY BANDING TOTAL
20 species
103 individuals

YEARLY BANDING TOTAL (2007)
58 species
1,259 individuals

26-YEAR BANDING GRAND TOTAL
(since 28 June 1982)
124 species
49,342 individuals

NOTABLE RECAPTURES THIS WEEK
(with original banding date, sex, and current age)

Carolina Chickadee (3)
06/29/05--3rd year male
07/13/05--3rd year male
07/16/06--2nd year female

White-throated Sparrow (3)
11/17/06--after hatch year unknown
01/14/07--after hatch year unknown
03/08/07--after hatch year male

Carolina Wren (1)
11/09/06--after hatch year female

OTHER NATURE NOTES OF INTEREST

--Although lasts week's year-high 23 species outpaced this week's 20, we banded more birds during the current period--103 vs. 62--thanks primarily to an influx of Yellow-rumped Warblers. Formerly known as Myrtle Warblers, yellow-rumps were quite common at Hilton Pond Center in the early 1990s, with 509 banded in the winter of 1991-92 and 263 in 1994-95. In the past 11 years, however, we've averaged only 24.7 yellow-rumps per winter. The decline is due in part to not being present to run mist nets at the Center during our winter hummingbird banding trips to Costa Rica, but over the past decade we've also seen many fewer yellow-rumps in the area on our annual York/Rock Hill (SC) Christmas Bird Count. This suggests that Yellow-rumped Warblers--like Purple Finches and Pine Siskins--may be facultative (not obligate) migrants that only come south when winter food in more northerly parts of their range becomes scarce.

--An enthusiastic group from South Carolina Wildlife Federation visited Hilton Pond Center this week for a Guided Field Trip. Among the most interesting things we found on the ground beneath a Pecan tree was a collection of twigs with leaves still attached--the work of Twig Girdler Beetles that had laid their eggs in the twigs and then weakened them so the small branches would fall to earth with beetle larvae inside. Soon thereafter the grubs would begin feeding on tree roots in underground chambers more protected from cold weather.

All text & photos © Hilton Pond Center


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