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THIS WEEK at HILTON POND
1-31 July 2021

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HILTON POND SUNSETS
(from our on-going series)

All text, maps, charts & photos © Hilton Pond Center

Sunset over Hilton Pond (above), 06 July 2021
Not many clouds this evening, but the Sun
filtering through the trees made for a
very curious reflection.



FOR GOODNESS SNAKE--
BEWILDERING RATSNAKE TAXONOMY
(PLUS MONOCHROMATIC WARBLERS)

You've heard the old story about how fishermen always exaggerate the size of their catch? Well, the same holds true for most folks who encounter a snake in the wild. Foot-long baby Garter Snakes become three-foot serpents in the observer's eye, and big wild blacksnakes magically grow to several yards in length.

All text, maps, charts & photos © Hilton Pond Center

Shortly after dawn on 6 July 2021 we encountered a truly large but docile Eastern Rat Snake (aka Black Rat Snake), Pantherophis (formerly Elaphe) obsoletus (photo above), nearly fully extended on a brushpile beside the old farmhouse at Hilton Pond Center. It was a couple of inches in diameter at its thickest part, and we estimated its length at more than six feet--which would make it a sizable representative of its species. (The documented record is an uncommonly long 8' 5".) It was certainly one of the largest snakes we've seen on the property during the past 40 years.

Not wishing to be accused of exaggeration, we decided after the glossy-black, white-chinned ratsnake moved on to try as best we could for an actual determination of its size. Since we had snapped a quick cell phone photo while the snake was stationary (see attached), we figured with a measuring tape we could do a point-to-point-to-point series of measurements, employing sticks and bark bits as our guides. Sure enough, with the tape we came up with a reading of at least 71"--not counting various kinks and undulations in the snake's body--to confirm our estimate of six-feet-plus.

The ratsnake was a perfect specimen with no visible scars. Since it's a constrictor, we're certain the serpent diligently spends its days (and nights) crawling (and climbing) to find and devour Hispid Cotton Rats, Eastern Chipmunks, and nesting birds at the Center. It's all part of the natural balance we try to maintain in a world that seems to become more and more unbalanced with every passing year.

All text, maps, charts & photos © Hilton Pond Center

Incidentally, you may have wondered about our word choice in the second paragraph above, and about all the parenthetical commentary. We wrote: "Eastern Rat Snake (aka Black Rat Snake), Pantherophis (formerly Elaphe) obsoletus." It seems the organism we all have known for generations as the Black Rat Snake, Elaphe obsoletus obsoletus (above) has gotten caught in the machinations of reptile taxonomists who have subjected the snake's chromosomes to expert analysis--and you KNOW what that means these days: Lumping and splitting!

"Lumping" refers to taking two or more organisms thought of as unique and re-classifying them into the same species, while "splitting" makes two or more species out of one. Traditionally, most of biological classification was based on an organism's external appearance and/or internal anatomy--and sometimes on reproductive behavior and/or geographic distribution. If two organisms looked pretty much alike and bred together they were in the same species, while if they were geographically disjunct or looked significantly different they were considered unique species.

All text, maps, charts & photos © Hilton Pond Center

And one other thing. As taxonomists look at new DNA evidence, they're also faced with re-examining taxonomy's historical record. Sometimes genetic studies reveal how fellow taxonomists along the way thought they were describing a new species when in reality that species wasn't really different from one described earlier. DNA studies--although technically complex--attempt to simplify matters by looking at an organism's genetic make-up and its true degree of relatedness to other organisms. In some cases this hasn't resulted in just "lumping" or "splitting" but to reshuffling taxonomic groups and even defining entirely ones. (In the bird world, for example, the Yellow-breasted Chat--long believed to be a Wood Warbler--is now alone in its very own family.)

So why was Elaphe changed to Pantherophis for our familiar backyard ratsnake at Hilton Pond Center? Originally, Elaphe was the genus used for all ratsnakes worldwide; all occur in the Northern Hemisphere and are slender non-venomous constrictors with squarish heads and flattened ventral surfaces. Anatomically and behaviorally--partly arboreal, for example (as in the photo above)--they are quite similar. However, when laboratory work showed European and Asian ratsnakes were genetically removed from our North American ratsnakes the groups were split, Elaphe remaining with the Old World species and Pantherophis being applied to those in the Western Hemisphere.

All text, maps, charts & photos © Hilton Pond Center

Pantherophis was first given to a North American ratsnake--the Corn Snake, Pantherophis guttatus (above)--by Austrian taxonomist Leopold Fitzinger in 1843, so modern taxonomists adopted that genus name for all the rest of the eastern North American ratsnakes. (Interestingly, the genus Elaphe was actually subject to a mega-split. Mitochondrial DNA work now calls for at least nine genera--including Pantherophis--for an abundance of ratsnakes originally lumped together as Elaphe spp.)

All text, maps, charts & photos © Hilton Pond Center
Yellow Ratsnake photo above courtesy Noah Mueller & Florida Museum

And all this brings to why what we have always called a Black Rat Snake is now being called Eastern Rat Snake--or maybe Gray Rat Snake or Texas Rat Snake or Everglades Rat Snake--all of which look different to the naked eye--as does the Yellow Rat Snake above. Yikes! What name should we use? The answer to that question depends on to whom you talk and/or which map you consult.

All text, maps, charts & photos © Hilton Pond Center

One map (above) from the 4th edition of Peterson Field Guide to Reptiles and Amphibians of Eastern and Central North America talks about just three species of ratsnakes in the East--Eastern, Western, and Gray. That would be all well and good, except for those cross-hatched "areas of uncertainty." On the South Carolina coast, for example, all Pantherophis individuals would be Eastern Rat Snakes, while here at Hilton Pond in the Piedmont they might be Eastern Rat Snakes OR Gray Rat Snakes--which despite their genes and names happen to LOOK very much alike. Will we need to run a DNA sample on every ratsnake at the Center to know for sure what it is? Theoretically, yes, but for practical reasons that's not going to happen.

(NOTE: To be honest, we always preferred the name "Pilot Blacksnake" to "Black Rat Snake," a title that arose from an old mountain myth that the species led rattlesnakes to safe winter denning sites. P.S. We also wonder whether "ratsnake" should be one word or two.)

All text, maps, charts & photos © Hilton Pond Center

A second even-more-detailed map (above) from the Ratsnake Foundation (yes, there is one) in part contradicts the first map and allows for five (possibly seven) subspecies of Pantherophis obsoletus--plus an entirely separate West Texas species called Baird's Rat Snake, P. bairdi. According to this map, any ratsnake slithering around Hilton Pond is good ol' Pantherophis obsoletus obsoletus, the Black (Eastern) Rat Snake, but what are the rest? Here are common names to match the subspecies on the map just above:

Pantherophis obsoletus lindheimeri, Texas Rat Snake
P. o. spilotes, Gray or Midland Rat Snake
P. o. quadrivittatus, Yellow Rat Snake
P. o. williamsi, Gulf Hammock Rat Snake
P. o. deckerti, Florida Keys Rat Snake
P. o. rossalleni, Everglades Rat Snake (below)

All text, maps, charts & photos © Hilton Pond Center
Everglades Rat Snakeling photo above courtesy Snakes At Sunset

In closing, if you find all this ratsnake lumping and splitting and nomenclature change to be complicated and confusing and frustrating you are not alone. Even herpetologists and ratsnake taxonomists can't agree on all the details we have outlined. For goodness snake, some of them still argue that Elaphe never should have been split (or lumped) in the first place. Others rail over Pantherophis being applied to any Black aka Eastern aka Gray Rat Snake that lays eggs and produces 10-inch snakelings like the one (at right) that hatched a few years back at Hilton Pond Center.

So who's right? We guess you'll just have to decide for yourself what to call these six-foot long native serpents--remembering all the while to base your preference on ratsnake scientific studies and not opinion.

All text, maps, charts & photos © Hilton Pond Center


A MONOCHROMATIC WARBLER

So let's move on from confusing ratsnake taxonomy to something less controversial: A little songbird that's been hanging around this summer at Hilton Pond Center. The bird of which we speak is the Black-and-white Warbler (BAWW, above), whose name pretty much describes its appearance. This month we observed up close just how monochromatic the species is when we captured three of them for banding.

All text, maps, charts & photos © Hilton Pond Center

On top of sporting white plumage with black highlights (or are they basic black with white markings?), BAWW are nore-or-less sexually monomorphic; i.e., the genders strongly resemble each other. The most noticeable difference is the darker black cheek of the adult male (see our file photo, above); this area is pale in females (as below), but immature males also tend to have lighter cheeks.

All text, maps, charts & photos © Hilton Pond Center

Since it was mid-July when we caught these three latest Black-and-white Warblers, it's reasonable to conclude they probably breed at or very near the Center--especially since one was an adult female with an active brood patch and the others were recent fledglings with short tails. Such evidence for local breeding is very strong, even though we've never found a BAWW nest on the property.

All text, maps, charts & photos © Hilton Pond Center
Black-and-white Warbler photo above courtesy Gretchen Locy

Black-and-white Warblers are among the 14 parulids that do breed in the Carolina Piedmont; the remaining "eastern" Wood Warblers--23 species--only pass through and nest at more distant locales from the North Carolina mountains north as far as Canada. (In the past 40 years 34 of the 37 eastern warblers have been encountered here at the Center; we're missing only Kirtlan's, Cerulean, and Mourning Warbler.) Likewise, the BAWW is one of even fewer warblers (just six) that might be seen around Hilton Pond during winter when the rest of their relatives are spending colder months in the Neotropics. (In addition to BAWW, our wintering species include Yellow-rumped Warbler, Pine Warbler, and Palm Warbler, plus the more elusive Common Yellowthroat and Orange-crowned Warbler.) A Black-and-white Warbler was spotted by Gretchen Locy on the December 2020 York-Rock Hill (SC) Christmas Bird Count--the first-ever for that event.

All text, maps, charts & photos © Hilton Pond Center



HILTON POND SUNSETS
(from our on-going series)

"Never trust a person too lazy to get up for sunrise
or too busy to watch the sunset."
--BHjr

All text, maps, charts & photos © Hilton Pond Center

Sunset over Hilton Pond (above), 21 July 2021
What you're looking at here is a hazy horizon--the result of
the first wave of smoke rolling into the Carolina Piedmont
from big wildfires in the northwestern U.S. It wasn't
heavy enough to block out the Sun by day or Moon
that evening, as happened in the Northeast.

Photoshop image post-processing for this page employs
DeNoise AI, Sharpen AI, and other Topaz Lab tools




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Don't forget to scroll down for Nature Notes & Photos,
plus lists of all birds banded or recaptured during the period.


"This Week at Hilton Pond" is written and photographed by Dr. Bill Hilton Jr., executive director of Hilton Pond Center for Piedmont Natural History

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Thanks to the following fine folks for recent gifts in support of Hilton Pond Center for Piedmont Natural History and/or Operation RubyThroat: The Hummingbird Project. Your tax-deductible contributions allow us, among other things, to continue writing, photographing, and sharing "This Week at Hilton Pond" with students, teachers, and the general public. Please see Support or scroll below if you'd like to make a gift of your own.

We're pleased folks are thinking about the work of the Center and making donations. Those listed below made contributions received during the period. Please join them if you can in coming weeks.

Gifts can be made via PayPal (funding@hiltonpond.org); credit card via Network for Good (see link below); or personal check (c/o Hilton Pond Center, 1432 DeVinney Road, York SC 29745). You can also donate through our Facebook fundraising page.

The following made contributions to Hilton Pond Center during the period 1-31 July 2021:

  • Anonymous $250 (via Network for Good)
  • Sara Brydges (repeat donor, via Network for Good)
  • Nina Cantillo (via payPal)
  • Amanda C. McNulty (repeat supporter; via Network for Good)
  • The following friends contributed via the "Donate" button on one of the Center's Facebook postings or fund-raisers; some may be repeat contributors. Several have set up through Facebook to make a recurring monthly donation to benefit the Center.
    Nick Dotson, Cherie Steele, Joe St. John, Krista Manus, Claire McConaughay, Iddling Babbler, Danny Raines, Shannon Steele, Linda Pelletier, Laura Clark, Maggie Norris, Liz Layton*, Russell Rogers, Gretchen Locy, Bill Pennington, Lynn Biasini McElfresh, Richard Barnett, Beatriz Moisset

    (
    * = past participant in Operation RubyThroat Neotropical Hummingbird expedition)
 
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BIRDS BANDED THIS WEEK at
HILTON POND CENTER
1-31 July 2021

SPECIES BANDED THIS PERIOD:
Ruby-throated Hummingbird--70
American Redstart--2
Carolina Chickadee--2
Black-and-white Warbler--3
White-eyed Vireo--2
American Goldfinch--1
Indigo Bunting--1
Great Crested Flycatcher--1
Northern Cardinal--3
Carolina Wren--3
House Finch--4
Tufted Titmouse--1
Mourning Dove--2

* = new banded species for 2021


PERIOD BANDING TOTAL:
13 species
95 individuals


2021 BANDING TOTAL:
55 species (40-yr. avg. = 63.7)

2,147 individuals
(40-yr. avg. =
1,848.8)

88 Ruby-throated Hummingbirds, 34 returns from previous years


40-YEAR BANDING GRAND TOTAL:
(Banding began 28 June 1982; since then 173 species have been observed on or over the property.)
127 species banded
73,950 individuals banded

6,732 Ruby-throated Hummingbirds banded since 1984

NOTABLE RECAPTURES THIS WEEK:
(with original banding date, verified sex, and current age):
Ruby-throated Hummingbird (19)
08/25/16--after 6th year female
08/01/17--5th year female
08/27/17--after 5th year female
06/24/19--after 3rd year female
07/06/19--3rd year female
07/27/19--3rd year female
08/25/19--after 3rd year female
08/29/19--3rd year female
09/13/19--3rd year female
06/25/20--after 2nd year female
06/26/20--after 2nd year female
07/05/20--after 2nd year female
07/08/20--after 2nd year female
07/15/20--2nd year female
07/22/20--2nd year female
07/29/20--2nd year male
07/30/20--after 2nd year male
07/31/20--2nd year female
09/15/20--2nd year male

Northern Cardinal (2)
10/15/17--5th year male
09/23/18--after 4th year male

Downy Woodpecker (1)
08/21/16--6th year male

House Finch (1)
03/18/18--4th year male

Tufted Titmouse (1)
11/08/20--2nd year male

* New longevity record for species
(none this week)

OTHER NATURE NOTES:
--Our oldest returning Ruby-throated Hummingbird (RTHU) in Jul 2021 was banded Aug 2016 and not encountered again at Hilton Pond Center until last year; now an after-6th-year bird, she likely was in the vicinity during the summers of 2017, 2018, and 2019 but we just didn't catch her. By comparison, two other female RTHU from 2017 have been recaptured every year since banding. (See complete recapture/return list at lower left.)

--Of 19 hummers recaptured this month from previous years, all but three were females, and no males were in the older years. This suggests either that males go somewhere else after being banded at the Center and/or they simply don't live as long.

--As of 31 Jul, the Hilton Pond 2021 Yard List stood at 79--about 46% of 172 avian species encountered locally since 1982. (Incidentally, 77 of those species so far this year have been observed from the windows or porches of our old farmhouse! Best year so far was 111 species in 2020. If you're not keeping a
Yard List for your own property we encourage you to do so, and to report your sightings via eBird,
where y
ou, too, can be a "citizen scientist!") New species observed locally during the period 1-31 Jul: Yellow-billed Cuckoo.

--Our immediate past installment of "This Week at Hilton Pond" was about yellows (and blacks) in nature. It's archived and always available on our Web site as Installment #746.

All text & photos © Hilton Pond Center



Please report your spring, summer &
fall sightings of
Color-marked
Ruby-throated Hummingbirds


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Hilton Pond Center for Piedmont Natural History is a non-profit research, conservation & education organization in York, South Carolina USA; phone (803) 684-5852. Directed by Dr. Bill Hilton Jr., aka "The Piedmont Naturalist," it is parent organization for Operation RubyThroat. Web site contents--including text and photos--may NOT be duplicated, modified, or used in any way except with express written permission of Hilton Pond Center. All rights reserved worldwide. To request permission for use or for further assistance, please contact Webmaster.